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RICINUS

Ricinus L., Sp. Pl. 2: 1007. 1753; Gen. Pl. ed. 5: 437. 1754; Muell. Arg. in DC., Prodr. 15(2): 1016. 1866; Benth. & Hook. f., Gen. Pl. 3(1): 321. 1880; Hook. f., Fl. Brit. Ind. 5: 457. 1890; Pax in Engl. Pflanzenreich 4. 147. 11: 119. 1919; Qiu & Gilbert, Fl. China @ eFloras.org 11: 248; Gillespie, Fl. North Amer. @ eFloras.org vol. 12; Radcliffe- Smith, Fl. Pak @ eFloras.org p. 69.

A large monoecious glabrous annual or perennial herb. Latex absent. Leaves alternate, petiolate, stipulate, simple, peltate, palmately-lobed, glandular-serrate at margin, penninerved. Petioles long, glandular at apex and base. Stipules united to form a caducous sheath, leaving circular scar at node. Inflorescence usually bisexual with proximal male flowers and distal female flowers or all flowers female, terminal or leaf-opposed subpaniculate racemes; bracts membranous, caducous, glands subtending each bract 2. Pedicels present, articulate. Male Flowers: Male buds globose. Calyx 3-5 lobed, valvate, closed in bud. Petals 0. Disc 0. Stamens numerous (up to 1000), with filaments variously united into about 25 slender, irregularly branched columns/ phalanges; anthers basifixed, bithecous, subglobose or globose, longitudinally dehiscent. Pistillode 0. Female Flowers: Pedicels extending in fruits; buds conical. Sepals 5, valvate, caducous. Petals 0. Disc 0. Ovary 3-locular with 1 ovule per loculus, usually echinate, occasionally smooth; styles 3, +/- free, bipartite, papillose-pumose, dark-red. Fruit a trilobate capsule, 3-locular, usually echinate, dehiscing into 3 bivalved cocci. Seeds dorsi-ventrally compressed, ovoid, smooth, carunculate, testa crustaceous.

One species

Ricinus communis

Ricinus communis L., Sp. Pl. 2: 1007. 1753; Muell. Arg. in DC., Prodr. 15(2): 1017. 1866; Hook. f., Fl. Brit. Ind. 5: 457. 1887; Parker, For. Fl. Punj. ed. 1: 461. 1918 (Reprint 1973); Pax in Engl. Pflanzenreich 4. 147. 11: 119. 1919; Fl. China @ eFloras.org 11: 248; Fl. North Amer. @ eFloras.org vol. 12; Radcliffe- Smith, Fl. Pak @ eFloras.org p. 69.

An evergreen, erect, soft-wooded monoecious shrub or occasionally a small tree, up to 3 m tall. Stem hollow. Younger parts glaucous, whole plant often reddish or purplish. Indumentum absent. Leaves alternate, petiolate, petiole stout, 18-45 cm long, hollow when mature, adaxially 2 sessile glands at apex and variable number (0-3) of different shapes present at/ near the base. Leaf blade circular in outline, 20-50 cm in diameter (near to size of petiole), peltate, palmately 7-12-lobed, medium lobe largest, 10-20 cm x 2-9 cm, the lateral ones progressively smaller, lobes lanceolate to oblanceolate, margin serrate or biserrate with irregular teeth, teeth gland-tipped, apex acute to acuminate; laminar glands scattered on adaxial surface. Stipules 2-3 cm long, connate forming sheath, sheath ovate, leaving a circular scar at node when fallen. Flowers large, in terminal 10-25 cm (35 cm in fruit) long subpanicle racemes, bisexual, the upper flowers female, lower ones male. Bracts 1 cm long. Male Flowers: Pedicels ca. 6 mm long, 3-5 partite, lobes 5-8 mm x 3-4 mm, slightly unequal, valvate, membranous, elliptic-ovate, acute, yellowish green, erect when young, reflexed at maturity. Petals 0. Disc 0. Stamens numerous up to 1000, with the filaments variously united into about 25 slender, irregular branched columns/ phalanges; stamen cluster +/- spheric, 10-12 mm diameter and 7-8 mm long; anthers ca. 0.4 mm long, pale yellow, basifixed, bithecous, globose or subglobose, longitudinal dehiscence. Pistillode 0. Female Flowers: Congested; pedicels 3-5 mm long, articulated, extending up to 4.5 cm in fruit. Calyx ca. 5 mm long, spathaceously splitting, lobes 5, ovate-lanceolate, acuminate, green-purplish, caducous. Disc 0. Staminodes 0. Ovary ca. 2 mm x 2 mm, trilobate-subglobose, densely covered with slender, conic, soft, bristle-tipped outgrowth, ovary 3-locular, 1 ovule per cell; styles 3, +/- free, 3-7 mm long, bipartite, papillose-plumose, dark red or orange-red. Fruit a trilobate capsule, ellipsoid or ovoid, 1.0-2.5 cm x 1-1.8 cm, 3- celled, echinate (sparingly to densely covered with narrowly cylindric bristle-tipped fleshy processes, 3-5 mm long), breaking into 3 2-valved crustacean cocci. Seeds 7-12 mm long, oblong, brown, mottled; caruncle depressed conical, 1-2 mm x 2-3 mm.

Flowering and Fruiting: June to October
Common Names: Castor Bean, Castor Oil Plant, Wonder Tree; Arand, Arandi (Hindi)

 Plant  Leaves  Young leaf with glands  Young leaf with apical,  basal petiolar glands and  marginal glands  Basal Petiolar Glands  Stipules  Inflorescence with distal female flowers and proximal male flowers  Inflorescence bearing Male Flowers below and Female Flowers above  Male Flower   Male Flower  Stamen clusters  Calyx in Male Flower  Female flowers  Female flower  T.S. Ovary Fruits Fruit with 3 cocci Seeds Seeds